A · A1 · A1b · A1b1 · BT · CT · CF · F · K · K2 · P · R · R1 · R1b · R-M343 · R-M269 · R-L23 · R-Z2103 · R-Z2106 · R-Z2109 · R-Z2110

Haplogroup R-Z2110

Macro-haplogroup
R
Parent clade
R-Z2109
Formed (estimate)
c. 4,000 - 4,500 years before present (estimate)
TMRCA (estimate)
c. 2,500 - 3,000 years ago (estimate)

Overview

Haplogroup R-Z2110 is a major downstream branch of R-Z2109 under the eastern R1b lineage R-Z2103. It likely arose in the later phases of the Early Bronze Age among steppe derived groups established in the Balkans and along the northern Aegean. Z2110 represents one of the principal lineages carrying Yamnaya related ancestry into southeastern Europe and into the emerging complex societies of the Balkans and Greece. The distribution of Z2110 and its subclades suggests that it participated in the same broad demographic processes that carried steppe ancestry into Thracian, Illyrian and early Greek speaking populations. Ancient DNA has revealed that steppe derived R-Z2103 lineages, including ancestors of Z2110, were common in Bronze Age individuals from Bulgaria, northern Greece and the lower Danube, indicating intense interaction between the steppe and the Balkan Peninsula. Today, Z2110 is represented by several downstream branches with distinct regional focuses in the Balkans and adjacent regions. Its internal topology points to both Bronze Age founder events and later historical movements, including the migrations and cultural transformations of the classical, Roman and medieval eras. As an integral part of the Z2103 complex, Z2110 is essential for understanding how eastern R1b contributed to the paternal structure of southeastern Europe.

Geographic distribution

Z2110 is most common in the Balkans, particularly in Bulgaria, Greece, Albania, North Macedonia and parts of Romania and Serbia. It is also present in Anatolia and the Caucasus at lower frequencies, and appears sporadically in Italy and central Europe.

Ancient DNA

  • Bronze Age individuals from Bulgaria and the lower Danube show Z2103 derived lineages whose structure is consistent with the emergence of branches like Z2110.
  • Ancient remains from Mycenaean and post Mycenaean Greece include R-Z2103 derived haplogroups that likely passed through nodes such as Z2110.
  • Iron Age and classical period populations in the Balkans and Aegean display continuing representation of eastern R1b lineages, including those under Z2110.
  • Patterns of continuity suggest that Z2110 bearing lineages remained rooted in southeastern Europe for more than three millennia.

Phylogeny & subclades

R-Z2110 is a key downstream branch of R-Z2109 and sits within the broader R-Z2106 cluster. It has several internal subclades that are concentrated in particular parts of the Balkans and Anatolia, reflecting region specific founder events.

  • R-Z2110*
  • Balkan regional microbranches
  • Aegean and Anatolian subclusters

Notes & context

Z2110 is one of the main representatives of steppe derived R1b in the Balkans and Aegean and is therefore critical for reconstructing Bronze Age and Iron Age demographic history in southeastern Europe.