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Haplogroup R-L295 (R2a1a)

Macro-haplogroup
R
Parent clade
R-P249
Formed (estimate)
c. 8800 - 9600 years before present
TMRCA (estimate)
c. 7000 - 8800 years ago

Overview

Haplogroup R-L295, also known as R2a1 in some classifications, represents one of the best documented downstream branches of R2a and provides an important window into early Holocene population movements across South Asia, the Near East and parts of Europe. It likely arose in a zone that connected South Central Asia with western Asia, at a time when farming economies were expanding and complex regional interaction networks were being established. R-L295 shows a wide but uneven distribution, with notable frequencies in South Asia and scattered occurrences in Anatolia, the Arabian Peninsula, Central Asia and Europe. This pattern suggests that its carriers were involved in multiple episodes of mobility, including both gradual range expansions of early farmers and later historical movements such as trade, military campaigns and religious migrations. Rather than a single rapid demographic expansion, L295 appears to have diversified through repeated, smaller scale founder events into different cultural and geographic contexts. From a phylogenetic perspective, L295 sits as a key subclade under R-P249 and is itself divided into additional branches, including R-L294 and various regionally restricted lineages. The age profile of L295 is compatible with its presence in Neolithic and early Bronze Age populations, although direct ancient DNA assignments remain limited. Its broad but low level presence across Eurasia makes it a valuable marker of long distance male mediated contacts linking South Asia with western Eurasian regions.

Geographic distribution

South Asia, Anatolia, the Arabian Peninsula, Central Asia and scattered occurrences in Europe and the Caucasus.

Ancient DNA

  • No securely assigned ancient individuals fall directly under L295, but several R2a related samples from the Near East and South Asia align with its upstream position.
  • The age estimates for L295 match the time frame of Neolithic and early Bronze Age expansions in South and West Asia.

Phylogeny & subclades

A major downstream branch of R-P249 within R2a, with internal division into L295* and further subclades such as R-L294.

  • R-L295*
  • R-L294

Notes & context

An important lineage connecting South Asian R2a diversity with more distant occurrences in western Eurasia.