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Haplogroup J2b-Z40091

Macro-haplogroup
J
Parent clade
J2b-M241
Formed (estimate)
c. 8,000 to 10,800 years before present
TMRCA (estimate)
c. 2,700 to 4,200 years ago

Overview

J2b-Z40091 is a downstream branch of J2b-M241 with a demographic center along the southern Balkans, the Aegean basin and western Anatolia. Its early formation corresponds to the spread of mid Holocene Balkan agricultural groups and the development of Aegean maritime interaction routes. Archaeological parallels include late Neolithic Balkan settlements, early Aegean island communities and proto urban coastal centers of western Anatolia. During the Bronze Age, Z40091 bearing groups were linked to Mycenaean expansions, Adriatic corridor exchanges and Aegean maritime circuits. Its downstream microstructure reflects coastal founder effects and demographic persistence into the Iron Age and classical eras, particularly in coastal Greek communities and western Anatolian seaboard settlements.

Geographic distribution

Common in Greece, Albania and western Turkey; moderate in Bulgaria, North Macedonia and Cyprus; lower in southern Italy and the Levant.

Ancient DNA

  • Balkan Chalcolithic individuals carry J2b structures aligning with early Z40091.
  • Aegean Bronze Age burials exhibit upstream forms of the lineage.
  • Western Anatolian Bronze Age individuals contain paternal markers tied to Z40091.
  • Iron Age Levantine and Aegean populations show downstream derivatives.
  • Classical Greek sites maintain continuity with multiple Z40091 branches.

Phylogeny & subclades

An Aegean and Balkan centered J2b cluster featuring several coastal microclades tied to maritime mobility.

  • Z40091*
  • Aegean coastal clusters
  • Western Anatolian derivatives

Notes & context

A major J2b lineage central to Aegean and southern Balkan Bronze Age population dynamics.