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Haplogroup J2a-Z7701

Macro-haplogroup
J
Parent clade
J2a-M67
Formed (estimate)
c. 10,000–13,000 ybp
TMRCA (estimate)
c. 6,000–8,500 ybp

Overview

J2a-Z7701 is a major early internal branch of the large and diverse Mediterranean-oriented J2a-M67 lineage. It represents a foundational upstream trunk from which several well-known downstream coastal, highland and Aegean-centered clades later emerged. Z7701’s temporal placement coincides with the expansion of advanced Neolithic agricultural economies and the earliest phases of long-distance maritime connectivity in the eastern Mediterranean. This makes Z7701 a key ancestral lineage linking Anatolia, Cyprus, the Levant and the Aegean in the early phases of farming dispersal and cultural interconnectivity.

Geographic distribution

Z7701 has a wide but regionally tiered geographic pattern. Highest frequencies occur across western and southwestern Anatolia, Cyprus, coastal Syria, Lebanon, Israel, Crete, the Cyclades, mainland Greece and parts of the central and southern Balkans. Additional distributions extend into southern Italy, Sicily, and occasionally Iberia. Inland, sporadic Z7701 lineages appear in Armenia, eastern Anatolia and the northern Fertile Crescent. Its distribution matches the spread of early ceramic Neolithic groups, later Bronze Age maritime polities and enduring Aegean cultural networks.

Ancient DNA

  • Early Aegean Neolithic individuals (e.g., Franchthi, Knossos) carry J2a-M67 branches compatible with Z7701’s upstream ancestry.
  • Kumtepe and early Troy phases exhibit J2a lineages associated with downstream Z7701 clusters.
  • Chalcolithic and Bronze Age Cypriot individuals show J2a phylogenies mapping to Z7701-derived Mediterranean subclades.
  • Late Bronze Age Levantine individuals from coastal trade hubs also contain M67/Z7701-related signatures.

Phylogeny & subclades

Z7701 sits below J2a-M67, one of the most expansive and structured branches of J2a. The clade occupies a top-level position in the M67 phylogeny from which multiple prominent maritime and Mediterranean-focused subclades descend. Its sister branches include Z500, M92 and other nodes forming the backbone of Aegean–Levantine J2a diversity. Z7701’s internal topology splits into highland, Aegean and Levantine groups, reflecting complex population interactions during the Neolithic, Chalcolithic and Bronze Age periods.

  • Z7701* basal maritime/Anatolian clusters
  • Z7701 > Aegean-centered lineages
  • Z7701 > Levantine coastal branches
  • Z7701 > Balkan and Mediterranean microclades

Notes & context

Z7701 is critical for understanding the demographic and cultural fabric of the eastern Mediterranean during the late Neolithic and Bronze Age. As a major parent node of downstream J2a-M67 coastal lineages, it provides structural context for interpreting Aegean, Cypriot, Anatolian and Levantine expansions.